diff --git a/data/templates/article.jats_publishing b/data/templates/article.jats_publishing
index 11902cae8..bb69404c2 100644
--- a/data/templates/article.jats_publishing
+++ b/data/templates/article.jats_publishing
@@ -81,6 +81,9 @@ $endif$
 $if(title)$
 <title-group>
 <article-title>$title$</article-title>
+$if(subtitle)$
+<subtitle>${subtitle}</subtitle>
+$endif$
 </title-group>
 $endif$
 $if(author)$
diff --git a/data/templates/default.jats_articleauthoring b/data/templates/default.jats_articleauthoring
index 66a4157d7..8cf89fa68 100644
--- a/data/templates/default.jats_articleauthoring
+++ b/data/templates/default.jats_articleauthoring
@@ -14,6 +14,9 @@ $endif$
 $if(title)$
 <title-group>
 <article-title>$title$</article-title>
+$if(subtitle)$
+<subtitle>${subtitle}</subtitle>
+$endif$
 </title-group>
 $endif$
 $if(author)$
diff --git a/doc/jats.md b/doc/jats.md
index bebb2536a..7eca75c8e 100644
--- a/doc/jats.md
+++ b/doc/jats.md
@@ -336,12 +336,21 @@ Metadata Values
 :   Additional notes concerning the whole article. Added to the
     article's frontmatter via the [`<notes>`][elem:notes] element.
 
+`subtitle`
+:   Subordinate part of the document title. Added to the
+    document's front matter as a
+    [`<subtitle>`][elem:article-title] element.
+
 `tags`
 :   list of keywords. Items are used as contents of the
     [`<kwd>`][elem:kwd] element; the elements are grouped in a
     [`<kwd-group>`][elem:kwd-group] with the
     [`kwd-group-type`][attr:kwd-group-type] value `author`.
 
+`title`
+:   The article title. Added to the document's front matter via the
+    [`<article-title>`][elem:article-title] element.
+
 Required Metadata
 -----------------
 
@@ -378,6 +387,7 @@ Required metadata values:
 [elem:abstract]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/abstract.html
 [elem:article-id]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/article-id.html
 [elem:article-meta]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/article-meta.html
+[elem:article-title]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/article-title.html
 [elem:copyright-holder]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/copyright-holder.html
 [elem:copyright-statement]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/copyright-statement.html
 [elem:copyright-year]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/copyright-year.html
@@ -400,6 +410,7 @@ Required metadata values:
 [elem:string-name]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/string-name.html
 [elem:subj-group]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/subj-group.html
 [elem:subject]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/subject.html
+[elem:subtitle]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/subtitle.html
 [elem:surname]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/surname.html
 [elem:xref]: https://jats.nlm.nih.gov/publishing/tag-library/1.2/element/xref.html